1.06i - Exponential growth and decay models

1.06i - Exponential growth and decay models

Build growth and decay models from equal proportional changes, fit their rates and find meaningful threshold times. Use half-lives, continuous compounding and nonzero baselines, then check predictions against observations and refine the assumptions when needed.

Constant proportional change

If a culture grows by the same percentage each month, how long before its size doubles? Start with 500500 organisms and a monthly multiplier e0.12\mathrm e^{0.12}. The first month's increase is about 6464, but the second month's increase is larger: the same percentage acts on a larger population. Equal additions would describe a different model.

You already know

eu+v=euev\mathrm e^{u+v}=\mathrm e^u\mathrm e^v, so adding to an exponent multiplies the output. To solve eu=v\mathrm e^u=v, with v>0v>0, use u=ln⁡vu=\ln v. Keep unrounded calculator values until the final answer.

A surprising consequence is that doubling the initial population does not shorten its doubling time. Both populations multiply by exactly the same factor over every equal interval. We will calculate the opening model's doubling time below.

Exponential growth and decay

An exponential model N(t)=N0ektN(t)=N_0\mathrm e^{kt}, with N0>0N_0>0, has a constant multiplicative change over equal time intervals. It grows when k>0k>0, decays when k<0k<0, and stays constant when k=0k=0. Use t≥0t\ge0 for time measured after a stated start, unless earlier times are justified.

Dividing two model values shows precisely which part stays constant.

Constant proportional change

For N(t)=N0ektN(t)=N_0\mathrm e^{kt},

N(t+h)N(t)=ekh.\frac{N(t+h)}{N(t)}=\mathrm e^{kh}.

The percentage change over one time unit is 100(ek−1)%100(\mathrm e^k-1)\%; for decay, the positive percentage decrease is 100(1−ek)%100(1-\mathrm e^k)\%. The known exponential gradient gives dN/dt=kN\mathrm dN/\mathrm dt=kN.

Not in the formula sheet: learn it.

The exponent must be dimensionless: if time is in months, kk has units per month. The instantaneous proportional rate kk is usually different from the fractional change over an entire month.

Worked example 1: interpreting growth

Question. For N=500e0.12tN=500\mathrm e^{0.12t}, with tt in months, find the initial population, the monthly percentage increase and the population after 33 months.

Plan. Set t=0t=0, interpret the one-month multiplier, then evaluate the requested time.

N(0)=500set the exponent to zero100(e0.12−1)≈12.7%convert multiplier to increaseN(3)=500e0.36≈716.665substitute the elapsed time\begin{aligned} N(0)&=500 &&\text{set the exponent to zero}\\ 100(\mathrm e^{0.12}-1)&\approx12.7\% &&\text{convert multiplier to increase}\\ N(3)&=500\mathrm e^{0.36}\approx716.665 &&\text{substitute the elapsed time} \end{aligned}

Why this step? A monthly multiplier 1.12749…1.12749\ldots means an increase of 0.12749…0.12749\ldots, not 1.12749…1.12749\ldots, of the previous amount.

Answer. Initially 500500; monthly growth about 12.7%12.7\%; about 717717 organisms after 33 months.

Check. The three-month multiplier is (e0.12)3=e0.36>1(\mathrm e^{0.12})^3=\mathrm e^{0.36}>1, consistent with growth.

Change only the sign of the rate constant. The amount still stays positive, but its multiplier is now less than 11.

Worked example 2: interpreting decay

Question. A count is modelled by N=500e−0.12tN=500\mathrm e^{-0.12t}, with tt in months. Find the monthly percentage decrease and the count after 33 months.

Plan. Subtract the multiplier from 11 to find the decrease, then evaluate the model.

q=e−0.12≈0.886920one-month multiplier100(1−q)≈11.3%fraction lost as a percentageN(3)=500e−0.36≈348.838three months of decay\begin{aligned} q&=\mathrm e^{-0.12}\approx0.886920 &&\text{one-month multiplier}\\ 100(1-q)&\approx11.3\% &&\text{fraction lost as a percentage}\\ N(3)&=500\mathrm e^{-0.36}\approx348.838 &&\text{three months of decay} \end{aligned}

Why this step? A negative continuous rate does not mean subtracting 12%12\% each month; exponentiate it first.

Answer. Monthly decrease about 11.3%11.3\%; a count of about 349349 after 33 months.

Check. 0<N(3)<5000<N(3)<500, as a positive decay model requires.

Population counts may be non-integer model outputs. Round to whole individuals when interpreting them, while retaining the unrounded outputs in calculations.

Watch out

For N=N0e0.12tN=N_0\mathrm e^{0.12t}, calling the one-unit increase exactly 12%12\% confuses a continuous rate with a whole-interval change. The increase is 100(e0.12−1)%100(\mathrm e^{0.12}-1)\%.

Use the multiplier to interpret a different decay rate.

Fitting rates and finding threshold times

Two positive observations determine the parameters of a chosen exponential model. They do not establish that an exponential is the right choice; reserve further observations for checking it.

Fitting an exponential rate

For N=N0ektN=N_0\mathrm e^{kt}, if N(t1)=N1>0N(t_1)=N_1>0, N(t2)=N2>0N(t_2)=N_2>0, and t2≠t1t_2\ne t_1,

N2N1=ek(t2−t1)k=ln⁡(N2/N1)t2−t1N0=N1e−kt1\begin{aligned} \frac{N_2}{N_1}&=\mathrm e^{k(t_2-t_1)}\\ k&=\frac{\ln(N_2/N_1)}{t_2-t_1}\\ N_0&=N_1\mathrm e^{-kt_1} \end{aligned}

Not in the formula sheet: learn it.

Division cancels the unknown initial coefficient. Use elapsed time t2−t1t_2-t_1, not whichever time label is larger.

Worked example 3: fitting from an initial observation

Question. An amount is 300300 at t=0t=0 and 450450 at t=4t=4 days. Assuming N=N0ektN=N_0\mathrm e^{kt}, determine the model and when it first reaches 600600.

Plan. Use the first observation for N0N_0, fit the rate from the ratio and solve the threshold equation.

N0=300the first time is zeroe4k=450/300=1.5use the second observationk=ln⁡1.54≈0.101366take logsekt=600/300=2set the target amountt=ln⁡2k≈6.84solve in days\begin{aligned} N_0&=300 &&\text{the first time is zero}\\ \mathrm e^{4k}&=450/300=1.5 &&\text{use the second observation}\\ k&=\frac{\ln1.5}{4}\approx0.101366 &&\text{take logs}\\ \mathrm e^{kt}&=600/300=2 &&\text{set the target amount}\\ t&=\frac{\ln2}{k}\approx6.84 &&\text{solve in days} \end{aligned}

Why this step? The target equation compares the target with the initial amount; the fitted rate must be kept unrounded.

Answer. N=300e(ln⁡1.5)t/4N=300\mathrm e^{(\ln1.5)t/4}, reaching 600600 after about 6.846.84 days.

Check. N(6)≈551<600N(6)\approx551<600 and N(7)≈610>600N(7)\approx610>600. If only whole-day observations count, the first qualifying observation is day 77.

Now shift only the observation times by two days. The ratio determines the same rate, but the coefficient at t=0t=0 is no longer the first observed amount.

Worked example 4: a different time origin

Question. The amount is 300300 at t=2t=2 and 450450 at t=6t=6 days. Determine the model and when it reaches 600600.

Plan. Fit the rate over the four-day interval, recover the value at zero and solve from the known time 22.

k=ln⁡(450/300)6−2=ln⁡1.54use elapsed timeN0=300e−2k≈244.949move back to time zero600=300ek(t−2)measure time from the known valuet=2+ln⁡2k≈8.84return to the stated clock\begin{aligned} k&=\frac{\ln(450/300)}{6-2}=\frac{\ln1.5}{4} &&\text{use elapsed time}\\ N_0&=300\mathrm e^{-2k}\approx244.949 &&\text{move back to time zero}\\ 600&=300\mathrm e^{k(t-2)} &&\text{measure time from the known value}\\ t&=2+\frac{\ln2}{k}\approx8.84 &&\text{return to the stated clock} \end{aligned}

Why this step? The coefficient of ekt\mathrm e^{kt} means the amount at time zero, not the amount in whichever observation happens to appear first.

Answer. N=300ek(t−2)N=300\mathrm e^{k(t-2)}, with k=ln⁡1.5/4k=\ln1.5/4; the threshold occurs at about day 8.848.84.

Check. Substituting t=2t=2 gives 300300, and t=6t=6 gives 300(1.5)=450300(1.5)=450.

A negative solution can describe a backward extension rather than a future event. For example, a growing model starting at 300300 does not fall to 200200 at a non-negative time. Check direction and domain before interpreting an answer.

Watch out

Do not round every threshold time upwards. Round up only when a discrete observation or completed-period rule asks for the first permitted integer time; otherwise report the continuous crossing at suitable accuracy.

Fit a decay model and find a later threshold.

Half-lives, doubling times and time constants

The initial amount cancels when you ask for a fixed multiplying factor. That is why equal half-lives remove equal fractions, rather than equal amounts.

Doubling time and half-life

For N=N0ektN=N_0\mathrm e^{kt}, with N0>0N_0>0,

D=ln⁡2k(k>0),H=ln⁡2∣k∣(k<0).D=\frac{\ln2}{k}\quad(k>0),\qquad H=\frac{\ln2}{|k|}\quad(k<0).

After jj half-lives the remaining fraction is 2−j2^{-j}. When k=0k=0, a nonzero constant amount has neither a finite doubling time nor a finite half-life.

Not in the formula sheet: learn it.

For example, 160e−(ln⁡2)t/6160\mathrm e^{-(\ln2)t/6} has a half-life of 66 hours: after 1818 hours, three halvings leave 160/8=20160/8=20. This simple ratio check is often faster than a fresh logarithmic calculation.

Worked example 5: doubling with a continuous rate

Question. For P=200e0.08tP=200\mathrm e^{0.08t}, with tt in years, determine its doubling time and its percentage increase over one year.

Plan. Set the ratio to 22, then separately calculate the annual multiplier.

2=e0.08Ddoubling ratioD=ln⁡20.08≈8.66take logs100(e0.08−1)≈8.33%annual increase\begin{aligned} 2&=\mathrm e^{0.08D} &&\text{doubling ratio}\\ D&=\frac{\ln2}{0.08}\approx8.66 &&\text{take logs}\\ 100(\mathrm e^{0.08}-1)&\approx8.33\% &&\text{annual increase} \end{aligned}

Why this step? A doubling time is determined by a ratio, so the starting amount 200200 cancels.

Answer. Doubling takes about 8.668.66 years; the annual increase is about 8.33%8.33\%.

Check. 200e0.08D=200(2)=400200\mathrm e^{0.08D}=200(2)=400, using the exact value of DD.

Change only the sign of the rate. We now seek halving, and both the time and the amount remain positive.

Worked example 6: halving with the opposite rate

Question. For P=200e−0.08tP=200\mathrm e^{-0.08t}, determine the half-life and the one-year percentage decrease.

Plan. Set the ratio to 1/21/2, then calculate the amount lost in one year.

12=e−0.08Hhalving ratioH=ln⁡(1/2)−0.08=ln⁡20.08≈8.66negative signs cancel100(1−e−0.08)≈7.69%annual decrease\begin{aligned} \tfrac12&=\mathrm e^{-0.08H} &&\text{halving ratio}\\ H&=\frac{\ln(1/2)}{-0.08}=\frac{\ln2}{0.08}\approx8.66 &&\text{negative signs cancel}\\ 100(1-\mathrm e^{-0.08})&\approx7.69\% &&\text{annual decrease} \end{aligned}

Why this step? A half-life is a positive elapsed time; dividing ln⁡2\ln2 by a negative signed rate without adjusting the sign would give an impossible negative half-life.

Answer. Half-life about 8.668.66 years; annual decrease about 7.69%7.69\%.

Check. After two half-lives the model gives 200/4=50200/4=50.

A decay model can instead be written N=N0e−t/τN=N_0\mathrm e^{-t/\tau}, defining its time constant τ=1/∣k∣\tau=1/|k|. At t=τt=\tau, the remaining fraction is e−1≈0.368\mathrm e^{-1}\approx0.368, not 1/21/2; its half-life is τln⁡2\tau\ln2. These time-constant relations are not in the formula sheet: derive them from the exponent.

Watch out

A time constant and a half-life are different. Test the stated time in the exponent: −1-1 leaves e−1\mathrm e^{-1}, whereas −ln⁡2-\ln2 leaves 1/21/2.

The opening culture's growth rate gives doubling time ln⁡2/0.12≈5.78\ln2/0.12\approx5.78 months. Test the corresponding fixed-ratio reasoning with decay instead.

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